Unit: Natural selection
Chapter: Hardy Weinberg equilibrium
Reference: Hardy-Weinberg principle, Importance
Learning objectives
- To describe the conditions under which allele and genotype frequencies will change in populations.
- To explain the impacts on the population if any of the conditions of Hardy-Weinberg are not met.
Hardy-Weinberg principle
- Coined by G. H. Hardy and W. Weinberg, which describes the genetic structure of evolving population.
- Mutations introduce new genes into species resulting a change in gene frequencies.
- Gene frequency is defined as the frequency with which a particular allele occurs in the population.
According to Hardy-Weinberg principle allele frequencies in a population are stable and is constant from generation to generation i.e., the gene pool remains constant, called genetic equilibrium.
According to this principle a population is only at equilibrium when there is-
- No mutation
- No natural or artificial selection
- No gene flow (no gene migration)
- No genetic drift
- No genetic recombination
Individual frequency for may be named as P, q. etc. in a diploid organism p and q represent the frequency of allele A and a. the frequency of AA individuals in a population is p2. This can be stated in another way, i.e., the probability that an allele A with a frequency of p appears on both the chromosome of a diploid individual is the product of the probabilities i.e., p2. Similarly, of aa is q2, of Aa is 2 pq. Thus, is a binomial expansion of (p+q)2. It is possible to calculate all allele and genotype frequencies using the expressions allele frequency p+q = 1 and genotype frequency p2+2pq+q2=1
Importance
For populations in the Hardy-Weinberg equilibrium, the equation connects allele frequencies to genotype frequencies. This equation allows us to compare a population's genetic structure over time to the expected genetic structure if the population were in Hardy-Weinberg equilibrium.
Solved examples
Example 1. If the frequency of non-tasters of PTC – Paper in a population is 16%, find out the frequency of dominant allele and the percentage of homozygous dominant individuals in the population.
Solution 1– The frequency of non-tasters (i.e., homozygous recessive genotype) is 16%,
this means that q2 = 16% = 16/100 = 0.16
Value of q = 0.16, = 0.4.
As p + q = 1, p = (1 – 0.4) = 0.6
Frequency of dominant allele (p) will, therefore, be 0.6 or 60%.
Now, the frequency of homozygous dominant genotype (p2) = 0.6 × 0.6 = 0.36, or 36%
Example 2. Which Hardy-Weinberg factor represents the frequency of heterozygous individuals in a population?
a) p2 b)2pq c) q2 d) p2 + 2pq
Solution 2. b. The heterozygous individuals in a population are 2pq
Summary
- Hardy-Weinberg principle is coined by G. H. Hardy and W. Weinberg, which describes the genetic structure of evolving population.
- According to Hardy-Weinberg principle allele frequencies in a population are stable and is constant from generation to generation i.e., the gene pool remains constant, called genetic equilibrium.
- The factors that affect the Hardy Weinberg principle includes gene flow, genetic drift, mutation, genetic recombination, and natural selection.